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For more information about this mode, see the Column Fill Mode section below. Column fill mode causes the visible columns in the control to fill the width of the control's display area.
The sizing mode for a column is actually determined by its Inherited Auto Size Mode property.
For example, if rows but not columns are configured for automatic sizing and Wrap Mode is enabled, users can drag column dividers to change the width of a column and row heights will automatically adjust so that cell contents are still fully displayed.